Doctoral / PHD Programs in Ecology, Evolution, Systematics and Population Biology

25 universities offer graduate PHD program in Ecology, Evolution, Systematics and Population Biology

Princeton University logo
Ranked as:  #1 in Best National University
Tuition:  $56,470 per year
Total Cost:  $112,940 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  New Jersey
Acceptance:  5.63%

Graduate study in the Ecology and Evolutionary Biology is designed to lead to the degree of Doctor of Philosophy (Ph.D.).

The special areas of strength in the department are evolution and genomics, ecology and the environment, behavior and organismal biology, conservation biology, and infectious disease. Graduate students also have excellent opportunities for combining several areas for innovative interdisciplinary work.

The graduate program is designed to develop both the breadth and depth of understanding that will enable graduates of the department to respond to future advances in the field. At the same time, students acquire the detailed knowledge and techniques needed to become effective scientists.

During the first two years, members of the committee recommend courses and are available for consultation on planning and initiating research projects.

Though there are few course requirements, all students must enroll in five departmental core courses in their first year. Graduate students may take additional advanced topic-related courses occasionally throughout their graduate career.

New students are encouraged to begin research projects as soon as possible. New students work with the DGS to identify a temporary research adviser and research project. In some cases, the initial project becomes the thesis topic, but many students work on several smaller projects before beginning the subject of their dissertation. Graduate students are expected to continue their research and training during the summer at Princeton, in the field, or at another laboratory.

The general examination consists of an oral examination, which typically lasts three hours. The committee may also include members of other departments within the University.

The Master of Arts (M.A.) degree is normally an incidental degree on the way to full Ph.D. candidacy and is earned after a student successfully passes the general examination. It may also be awarded to students who, for various reasons, leave the Ph.D. program, provided that this requirement has been met.

All graduate students are required to teach at least two terms during the first four years of the program. Students who are not supported by an external fellowship, (e.g., NSF, NSERC, etc.) must teach an additional two terms.

In order to complete the Ph.D., a student consults with the committee members who evaluate the student dissertation. Once the committee approves the dissertation, the candidate has two weeks to prepare and present to the department a defense of the work in the final public oral examination for the doctoral degree.

Permanent courses may be offered by the department or program on an ongoing basis, depending on curricular needs, scheduling requirements, and student interest. Also not listed are graduate-level independent reading and research courses, which may be approved by the Graduate School for individual students.

EEB 502 Fundamental Concepts in Ecology, Evolution, and Behavior I.

Advanced discussions of ecology, evolution, and behavior focus on 50 fundamental papers. Ecological topics include dynamics and structure of populations, communities, ecosystems, and conservation biology. Behavioral topics include instinct and learning, social behavior, physiological ecology, and the evolution of behavior. Evolutionary topics include speciation, evolutionary and quantitative genetics, molecular evolution, evolutionary stable strategies, and evolution of development. (These are core courses).

EEB 504 Fundamental Concepts in Ecology, Evolution, and Behavior II.

Advanced discussions of ecology, evolution, and behavior focus on 50 fundamental papers. This course continues the discussion of EEB 502, q.v. The two courses are offered in alternate years, and may be taken in either order. (This is a core course.).

This course will cover the essential topics of what constitutes responsible conduct in research.

EEB 507 Recent Research in Population Biology.

Systematic reviews of recent literature in areas of ecology, evolution, and animal behavior are made. The general survey of literature is supplemented with detailed discussion of selected research papers of unusual importance and significance. (This is a core course.).

Intensive three-week field course undertaken in January or February in a suitable tropical locality. There are readings, discussions, and individual projects. The content and location are varied to suit the needs of the participants. Students provide their own travel funds. (This is a core course.).

EEB 522 Colloquium on the Biology of Populations.

Discussion of the central problems of population biology and approaches that have proved fruitful.

Discussion of current topics in evolutionary biology, ranging from historical processes inferred from the fossil record to microevolution demonstrated by observation and experiment. Topics vary from year to year, depending on current controversial issues.

This half-term topics course offers students exposure to current issues in conservation across the globe.

Discussion of the growing points in population and community ecology is held. Topics vary from year to year and include such subjects as the regulation of population numbers, the organization of communities, conservation, and the variation in ecological processes at different spatial and temporal scales.

Close reading of published papers illustrating the principles, achievements and difficulties that lie at the interface of theory and experiment in biology. Two important papers, read in advance by all students, will be considered each week emphasis will be on student discussion, not formal lectures. Topics include: cooperativity, robust adaptation, kinetic proofreading, sequence analysis, clustering, phylogenetics, analysis of fluctuations, maximum likelihood methods.

These are courses intended to help students develop and apply skills in the application of scientific, technological, and environmental analyses to problems of policy interest. Fall courses are numbered 585, Spring courses are numbered 586.

˅ More Details

Ecology and Evolutionary Biology - Graduate School

  • GRE Required:  Yes
  • Research Assistantships:  733
  • Teaching Assistantships:  655
  • Financial Aid: Register to view the details
Harvard University logo
Ranked as:  #3 in Best National University
Tuition:  $50,654 per year
Total Cost:  $101,308 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  Massachusetts
Acceptance:  5.01%

Alongside our core questions related to what makes humans unique, many members of our department are also interested in why and how our unique evolutionary history matters for practical and policy issues. Key questions revolve around health, exercise, immunity, innovation, and the construction of effective institutions.

In our research, we integrate insights on contemporary biology, physiology, genetics, anatomy and behavior from around the globe and well back into our evolutionary past. In building our community of scientists, including our students, we actively seek to expand our diversity, seeking people from different backgrounds, countries, cultures, experience, and training. It is at the heart of our approach to understand the human condition and all of its complex variation.

HEB is proud to continue a century-long tradition of training the future leaders in understanding humans from an evolutionary perspective. Our graduate students receive generous funding for their entire degree program and get hands-on training in teaching and both laboratory and field-based research. In addition to the extraordinary resources within the department, HEB graduate students also benefit from the unparalleled resources of Harvard University, including strong collaborations with departments such as Organismic and Evolutionary Biology, Economics and Psychology as well as the Broad Institute, Harvard Medical School, Kennedy School and the Peabody Museum.

Our interdisciplinary approach thus includes field and laboratory programs in many sub-disciplines including:.

The first two years are a combination of classwork and research. Ordinarily, students define a PhD topic in the third year, and then finish by the sixth year.

All HEB students receive five years of full funding, including tuition and stipend plus substantial departmental support for research. Training to teach is also an important component of the PhD.

˅ More Details

Human Evolutionary Biology

  • GRE Required:  Yes
  • Research Assistantships:  864
  • Teaching Assistantships:  1388
  • Financial Aid: Register to view the details
Brown University logo
Ranked as:  #13 in Best National University
Tuition:  $60,363 per year
Total Cost:  $120,726 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  Rhode Island
Acceptance:  7.67%

The graduate program in Ecology, Evolution, and Organismal Biology is intended for highly qualified students who plan to pursue a career that includes research in the fields of ecology and environmental science, organismal biology and or evolutionary biology.

Independent research begins early in the program, which is designed to prepare students for careers in research and higher education. Each student develops a program of preparation based on close mentoring by a thesis committee.

˅ More Details

Ecology, Evolution, and Organismal Biology

  • GRE Required:  Yes
  • Research Assistantships:  507
  • Teaching Assistantships:  483
  • Financial Aid: Register to view the details
we can find best-fit college

69 universities offer the Master's program in Ecology, Evolution, Systematics and Population Biology.

Which one best suits your need?

Please select a program.
Select a program:
Please choose a course.
Please choose a specialization.
Please choose an option.
Have you taken the GRE?
GRE quantitative field is required. (Minimum: 80 | Maximum: 170)
GRE Verbal field is required. (Minimum: 80 | Maximum: 170)
GRE AWA field is required. (Minimum: 0 | Maximum: 6)
Please choose a type of university.
Type of University:
Education level completed:
Please choose a price interval.
$5000 $80000
Tuition sorting:
Please provide a First Name.
Please provide a Last Name.
Please provide a name.
Please provide your address.
Please, enter valid email address.
Please, enter valid phone.
Please, enter valid phone.
Rice University logo
Ranked as:  #15 in Best National University
Tuition:  $47,913 per year
Total Cost:  $95,826 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  Texas
Acceptance:  10.89%

Doctor of Philosophy (PhD) Degree in the field of Ecology and Evolutionary Biology.

Program Learning Outcomes for the PhD Degree in the field of Ecology and Evolutionary Biology.

Upon completing the PhD degree in the field of Ecology and Evolutionary Biology, students will be able to:.

Demonstrate an ability to understand and critically evaluate concepts, research accomplishments, and techniques in ecology and evolutionary biology.

Demonstrate independent problem solving and critical thinking skills by identifying novel research questions in ecology and evolutionary biology and synthesizing critical paths towards answering them.

Demonstrate technical proficiency in a range of ecology and evolutionary biology research methods.

Requirements for the PhD Degree in the field of Ecology and Evolutionary Biology.

Most of the formal course studies will be completed in the first year of residence to allow the students to begin thesis research at the end of their second semester at Rice. Students should have completed coursework in ecology, evolution (or equivalent), mathematics (including calculus), and statistics prior to admission.

The requirements listed in the General Announcements (GA) satisfy the minimum requirements for this degree program.

Total Credit Hours Required for the PhD Degree in the field of Ecology and Evolutionary Biology.

CORE COURSE IN ECOLOGY AND EVOLUTIONARY BIOLOGY (course repeatable for credit).

GRADUATE SEMINAR IN ECOLOGY AND EVOLUTIONARY BIOLOGY (required in all years of residency, fall semester).

GRADUATE SEMINAR IN ECOLOGY AND EVOLUTIONARY BIOLOGY (required in all years of residency, spring semester).

GRADUATE TEACHING IN ECOLOGY AND EVOLUTIONARY BIOLOGY (two semesters) 1.

Select 2 courses from the following (2 semesters of any combination of BIOS Topics courses):3.

BIOS 801Ecology Evolutionary Biology Graduate Research credit hours vary depending on the number of other courses the student is taking in a given semester.

At least two topics courses must be completed before candidacy. Students are strongly encouraged to take at least one topics course per semester during all years of residency.

Students must maintain a minimum grade average of B (3.00 grade points) in courses taken in the department and satisfactory grades in courses taken outside the department. The review process requires that each student:.

Presents a public seminar on their research at the annual EEB Graduate Student Symposium.

Completing an original investigation and a doctoral thesis with at least three chapters with the potential to produce publications in reputable, peer-reviewed scientific journals.

Policies for the PhD Degree in the field of Ecology and Evolutionary Biology.

Applicants for graduate study in the Ecology and Evolutionary Biology Program must have:.

BA or BS degree or equivalent that provides a strong background in biology.

Strong ability and motivation, as indicated by academic record and recommendations.

Scores from the GRE biology subject exam are optional but can be helpful, particularly for students with nontraditional backgrounds in biology.

These requirements do not preclude admission of qualified applicants who have majored in areas other than biology. Although the program offers MS degrees, only on rare occasions are students who do not intend to pursue the PhD admitted to the graduate program. For general university requirements, see Graduate Degrees.

Some departments and programs have additional restrictions on transfer credit.

Students pursuing the PhD degree in the field of Ecology and Evolutionary Biology should be aware of the following departmental transfer credit guidelines:.

Opportunities for the PhD Degree in the field of Ecology and Evolutionary Biology.

All full-time Ecology and Evolutionary Biology graduate students receive funding and full tuition waivers as specified in their offer letters. Information Student Resources, Attendance at Scientific Conferences, Internships, Graduate Students Awards, the Graduate Student Association, etc.

˅ More Details

Doctor of Philosophy (PhD) Degree in the field of Ecology and Evolutionary Biology Rice

  • GRE Required:  Register to view the details
  • Research Assistantships:  Register to view the details
  • Teaching Assistantships:  Register to view the details
  • Financial Aid: Register to view the details
Cornell University logo
Ranked as:  #17 in Best National University
Tuition:  $30,042 per year
Total Cost:  $60,084 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  New York
Acceptance:  10.71%

The program is intended to provide students with broad exposure to concepts and research approaches within ecology and evolution (primarily through seminars and formal course work), as well as in-depth study in one or subdisciplines (normally guided by the student Special Committee). The specific course of study is tailored to the needs of the individual student. Excellent laboratory and nearby field facilities are available, as are numerous collections and libraries. All students are required to obtain teaching experience by serving as teaching assistants for at least two semesters.

All Graduate School Requirements, including the TOEFL Exam or IELTS Academic Exam for non-native English applicants.

Note: GRE test scores are no longer required for application to this program.

Please submit a combined Academic and Personal Statement of Purpose that outlines your reasons for pursuing graduate research and explains your academic interests and your broader background, experiences, and skills that can lead to a successful graduate school experience. This Statement of Purpose should provide the admissions committee with a sense of you as a whole person, and provide insight into your potential to contribute positively to a diverse and inclusive community.

Please describe how your background and experiences influenced your decision to pursue a graduate degree.

(2) Academic background and preparation (400 words): Describe your academic training, skills, research experience, and accomplishments relevant to your future graduate work. You may also provide the context around any perceived gaps or weaknesses in your academic record.

4) Importance of community, diversity, and inclusion (400 words): We strive to build a diverse and inclusive community that strengthens our intellectual and collaborative department. Please provide insight into your potential to contribute to a community of inclusion, belonging, and respect where scholars representing diverse backgrounds, perspectives, abilities, and experiences can learn and work productively and positively together.

Within sections 1 and 4 of the Statement of Purpose, you may also include relevant information on any of the following:.

How your personal, academic, and or professional experiences demonstrate your ability to be both persistent and resilient especially when navigating challenging circumstances.

How you have experienced or come to understand the barriers faced by others whose experiences and backgrounds may differ from your own.

A candidate for a doctoral degree in Ecology and Evolutionary Biology is expected to demonstrate a broad based knowledge in chosen discipline (Ecology, Evolutionary or Organismal Biology, or some combination), mastery of knowledge in the chosen sub-discipline (e.g., Ecosystem Biology, Community Population Ecology, Evolutionary Genetics, Macroevolution, Systematics, Molecular Evolution, or some combination), and create new knowledge, making an original and substantial contribution to the sub-discipline in a timely fashion.

Demonstrate broad-based knowledge in the discipline of Ecology, Evolutionary Biology, Organismal Biology, or some combination.

Be knowledgeable of organisms and ecological or evolutionary systems pertinent to doctoral research.

Effective teaching skills in ecology and evolutionary biology.

Involvement in professional activities and organizations related to academic discipline.

Cornell NYC Tech Distance, Hybrid, or Online Option Geneva Ithaca.

˅ More Details

Graduate School

  • GRE Required:  Register to view the details
  • Research Assistantships:  Register to view the details
  • Teaching Assistantships:  Register to view the details
  • Financial Aid: Register to view the details
Columbia University in the City of New York logo
Ranked as:  #18 in Best National University
Tuition:  $51,194 per year
Total Cost:  $102,388 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  New York
Acceptance:  6.66%

The Ecology, Evolution, and Environmental Biology (E3B) at Columbia University was established in 2001. In creating E3B, the university recognized that the fields of ecology, evolutionary biology, and environmental biology constitute a distinct subdivision of the biological sciences with its own set of intellectual foci, theoretical foundations, scales of analysis, and methodologies.

Our educational programs emphasize a multidisciplinary perspective to understand life on Earth from the level of organisms to global processes that sustain humanity and all life.

Through this collaboration, the department is able to tap into a broad array of scientific and intellectual resources in the greater New York City area.

Fellowships are awarded for those admitted to the PhD program in recognition of academic achievement and in expectation of scholarly success. Teaching and research experience are considered an important aspect of the training of graduate students. Thus, graduate fellowships include some teaching and research apprenticeship.

It is also desirable that students have had course work in calculus, physics, chemistry, statistics, genetics, ecology, evolution, and other areas of organismal biology. Prior research experience in a relevant area is strongly recommended. Students are admitted to begin only in the fall semester.

Columbia has offered a PhD specialization in Evolutionary Primatology for over two decades as part of the New York Consortium in Evolutionary Primatology (NYCEP). NYCEP, a consortium of the City University of New York, Columbia University, New York University, the American Museum of Natural History, and the Wildlife Conservation Society, provides a multi-institutional venue for doctoral-level training, which emphasizes all aspects of the behavioral, morphological, and evolutionary biology of primates. Course offerings in this program are coordinated across the NYCEP institutions.

˅ More Details

Ecology, Evolution, and Environmental Biology, PhD

  • GRE Required:  Register to view the details
  • Research Assistantships:  Register to view the details
  • Teaching Assistantships:  Register to view the details
  • Financial Aid: Register to view the details
University of California-Los Angeles logo
Ranked as:  #20 in Best National University
Tuition:  $28,131 per year
Total Cost:  $56,262 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  California
Acceptance:  14.33%

Ecology and Evolutionary Biology.

The doctoral program prepares students to excel in research, academia, industry, and governmental non-governmental organizations. Students have access to seminars and courses on a wide range of topics in evolutionary biology, ecology, and animal behavior, as well as biomathematics, theoretical modeling, and statistics. After completing a common set of course requirements during their first year, doctoral students focus on their research in one of our specializations.

Incoming doctoral students receive five-year support packages that typically include a combination of support in the form of fellowships, graduate student researcher appointments, and teaching assistantships. Many of our continuing students are successful at earning competitive campus and extramural graduate fellowships.

First complete and submit the UCLA Online Graduate School Application by the December 1st deadline. You will be asked to create an account, and we strongly recommend writing down your password. Select the Fall 2023 admission term, and Biology PhD.

You will need the following information to complete the online application:.

Ecology and Evolutionary Biology Supplementary Essay (800 word limit): The essay should tell the admission committee your background in Ecology and Evolutionary Biology and proposed focus, and give the committee information your work and professional experience. The essay should address the following:.

Our department in particular is looking for applicants with knowledge and creativity in their research area of interest, strong communication skills, resiliency in the face of setbacks, capacity for planning, organization, and teamwork, and high standards of ethics and integrity. We encourage applicants to share examples that highlight strengths in these areas.

Personal History Statement (500 word limit): The Personal Statement is an opportunity for you to provide additional information that may aid the selection committee in evaluating your preparation and aptitude for graduate study at UCLA. Please respond to one or of the following prompts. To be considered for a Cota-Robles fellowship, be sure to describe your contributions to diversity. The University of California Diversity Statement can be found online here.

This is a mandatory university-wide prompt for all graduate programs. (1500 word limit).

Names and email addresses of three recommenders. Specific instructions concerning letters of recommendation are described in the online application.

Awards and other Distinctions. Any academic awards, prizes, honors, fellowships or other distinctions that you have received.

ONE official, sealed copy of your academic transcripts.

Please note GRE scores are no longer considered during the application review process. You may see a place for them on the general application, but do not have an official score report sent to our department.

The Ecology and Evolutionary Biology PhD Program is one of 18 Graduate Programs in the biosciences.

MS Biology studying the Ecology and Evolution of Medicine.

˅ More Details

Ecology and Evolutionary Biology

  • GRE Required:  Register to view the details
  • Research Assistantships:  Register to view the details
  • Teaching Assistantships:  Register to view the details
  • Financial Aid: Register to view the details
University of Michigan-Ann Arbor logo
Ranked as:  #25 in Best National University
Tuition:  $49,548 per year
Total Cost:  $99,096 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  Michigan
Acceptance:  26.11%

U-M Gateway feature: Protecting algae-eating fish insufficient to save imperiled coral reefs Published online in the journal Nature Ecology Evolution by Jake Allgeier and Timothy Cline.

The Ecology and Evolutionary Biology is delighted to present the EEB First Paper Award to recognize students for their first lead-authored publications.

Cover story in Science: termites may have a larger role in future ecosystems.

EEB Student Dissertation Defense: Toxic Forms Most Beautiful: The Evolutionary Dynamics of Rear-Fanged Snake Venoms.

Peter Cerda, PhD Student, Ecology and Evolutionary Biology.

EEB Thursday Seminar Hybrid: #Findthatlizard: ecology, science outreach, and supporting underrepresented groups in herpetology and natural resources.

˅ More Details

Ecology and Evolutionary Biology

  • GRE Required:  Register to view the details
  • Research Assistantships:  Register to view the details
  • Teaching Assistantships:  Register to view the details
  • Financial Aid: Register to view the details
University of California-Davis logo
Ranked as:  #38 in Best National University
Tuition:  $28,700 per year
Total Cost:  $57,400 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  California
Acceptance:  46.35%

Research and training programs funded by the U.S. National Science Foundation, such as Graduate Research Fellowship Program (GRFP) and Integrative Graduate Education and Research Traineeships (IGERT), offer short-term support to the Center graduate student affiliates (both students in the Population Biology and in other UC Davis graduate groups), and provide support for independent postdoctoral associates who have recently completed their doctoral dissertations.

˅ More Details

Center for Population Biology

  • GRE Required:  Register to view the details
  • Research Assistantships:  Register to view the details
  • Teaching Assistantships:  Register to view the details
  • Financial Aid: Register to view the details
Ohio State University-Main Campus logo
Ranked as:  #49 in Best National University
Tuition:  $37,141 per year
Total Cost:  $74,282 * This tuition data is based on IPEDS. For the latest tuition amount, refer to the respective college websites.
State:  Ohio
Acceptance:  68.49%

A Study with Captive Populations.

New Tools for Quantifying Textbook Design for Biology Education.

The Evolution, Ecology, and Organismal Biology (EEOB) offers M.Sc. and Ph.D. degrees, as described in the EEOB Graduate Program Handbook.

We provide PhD and MS students with the skills needed to conduct original research and qualify for rewarding careers in our discipline. EEOB encompasses the study of animals, plants, and microbes in their relationships among themselves and their environments, and in their common ancestry. Our successful graduate students have found employment in academia, in state and federal agencies, and in ecological life science organizations and businesses across the country. EEOB knows that diversity in our ranks improves all aspects of our professional efforts. As a result, we are committed to recruiting, admitting and mentoring students from diverse backgrounds.

Benefits of Completing a Degree in the EEOB Graduate Program.

The Ohio State University is an R1 research institution, with ample resources and opportunities available to EEOB graduate students.

Most of our graduate students are funded by teaching or research assistantships (20 hours per week), or sometimes by University Fellowships, at the rate of ~$2,400 per month, plus a full tuition waiver and employer supported benefits.

The deadline for applications is November 15th, with admission starting in Autumn Semester of the following academic year. We no longer require GRE scores for admission to our program they may be submitted, if you like.

˅ More Details

Evolution, Ecology and Organismal Biology

  • GRE Required:  Register to view the details
  • Research Assistantships:  Register to view the details
  • Teaching Assistantships:  Register to view the details
  • Financial Aid: Register to view the details

What kind of scholarships are available for Graduate Programs in Ecology, Evolution, Systematics and Population Biology?

We have 154 scholarships awarding up to $1,256,696 for Masters program in for Ecology, Evolution, Systematics and Population Biology, targeting diverse candidates and not restricted to state or school-based programs.

Scholarship nameAmountCredibility
George and Lavinia Blick Research Fund$25,000High
STEM Teacher Graduate Scholarships$2,500High
PCI Women in STEM Scholarship$2,000High
ABC Humane Wildlife Women In STEM Academic Scholarship$1,000High
GMiS STEM Scholarships$500High

Find scholarships and financial aid for Ecology, Evolution, Systematics and Population Biology graduate programs

$500 $20000

How can I compare the Ecology, Evolution, Systematics and Population Biology Graduate Programs?

Compare the GRE score requirements, admission details, credit requirements and tuition for the Master's Program, from 69 universities offering Graduate PHD/Doctoral Programs in Ecology, Evolution, Systematics and Population Biology. Compare Graduate PHD/Doctoral Programs in Ecology, Evolution, Systematics and Population Biology

How will you rate the search results?

 0.0 rating, based on 0 reviews

Your review has been sent

Select rating
Your review must be at least 10 characters long
Your review must be not longer than 2000 characters